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In particular, Dr Lönnberg describes the skulls of two stags killed in Glenquoich Forest in Invernesshire, north-west Scotland.

The skulls displayed some features in common with Swedish (It is accordingly neither identical with the typical race of southern Sweden nor with the race of western Norway and most probably forms an independent geographic race or subspecies which suitably may be termed .

In 1806 Pennsylvanian-born naturalist and physician Benjamin Smith Barton suggested that North American elk and Red deer from Europe were sufficiently different to be considered different species and proposed the name wapiti, meaning “white rump”, for the North American elk.

Since then, the wapiti has been the subject of much taxonomic yo-yoing, being moved between a full species, ).

Generally-speaking, it is considered that in order for two individuals to be considered for subspecific status, there should be a maximum of 10% overlap in physical characteristics – in other words, they should be at least 90% different from each other.

Drs Lowe and Gardiner examined the skulls of 10 of the 19 subspecies of Red deer listed by John Ellerman and Sir Charles Morrison-Scott in their 1951 , examining 16 variables of skull size and shape and subjecting the data to three separate statistical analyses.

In the paper Dr Lönnberg compared the skull anatomy of Red deer collected from various parts of its range and proposed several of the 12-or-so subspecies still in contention today.

The wapiti range over much of North America and eastern Asia and are superficially similar to the Red deer of Europe and Asia (an area collectively termed “Eurasia”).